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Nature 552 (7684), 194199

[5] Chen CR, Pichurin P, Nagayama Y, Latrofa F, Rapoport B, McLachlan SM

Additionally, continued exposure of T cells at sites of inflammation elicits high levels of reactive oxygen species (ROS), which results decreased intracellular levels of GSH, dysregulation of redox balance, impaired signal transduction of the TCR/CD3 complex, and ultimately to synovial T cell hyporesponsiveness during RA [137]