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A 2012 paper in Oxidative Medicine and Cellular Longevity confirmed that GHK-Cus biological activity depends on proper copper complexation

Application of exogenous LOXL2 to cultured cartilage explants (derived from bovine articular cartilage) and engineered chondrogenic constructs (primary chondrocytes cultured to create a self-assembled structure), which were later implanted into athymic mice, promoted tissue maturation (increased mature crosslinks) and enhanced tensile mechanical properties (Makris et al ., 2014)

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[DOI] [PubMed] [Google Scholar] 64.Ignatov A., Lintzel J., Kreienkamp H.J., Schaller H.C