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253 UPR consists of three transmembrane proteins: protein kinase RNA-like ER kinase (PERK), 254 activating transcription factor 6 (ATF6), 255 and inositol-requiring signaling protein-1 (IRE1), 256 which all form stable complexes with the regulatory protein glucose regulatory protein 78 (GRP78, also known as Bip) under normal conditions, while upon ER stress, they dissociate from GRP78 and activate downstream signaling pathways