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Retinol works in several ways

10.1038/nature09486 Nature 132 LautenschgerJ.SchierleG

10.1016/j.xpro.2025.103863 22 ChaoulV.DibE

The peptides acidic residues (glutamate at position 2 and aspartate at position 3) have been proposed to facilitate interactions with chromatin-associated proteins and transcription regulatory complexes, consistent with observed epigenetic effects documented in in-vitro model systems